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Published In: Bryologia Europaea 3: 110. 236 (fasc. 25–28 Mon. 8. 1). 1845. (Bryol. Eur.) Name publication detail

Project Name Data (Last Modified On 3/16/2009)
Acceptance : Accepted
Project Data     (Last Modified On 5/4/2009)
General Reference:

Illustration & Map     (Last Modified On 5/4/2009)
Illustrations: Grimmia anodon Bruch & Schimp. (Figs. 1–7.)
Map: Grimmia anodon Bruch & Schimp. (Distribution in China)

Distribution     (Last Modified On 5/4/2009)
Habitat: on dry, exposed limestone or calcareous sandstone; alt. 1100–4850 m.
Distribution: China, India, Pakistan, Turkey, Armenia, Kazakstan, Russia, Europe, Africa, North and South America.

Specimens Examined     (Last Modified On 5/4/2009)
Specimens Examined:
Chinese specimens examined: Qinghai: Nang-qian Co., B. C. Tan 95-1705 (MO). Xinjiang: He-jiang Co., Z.-L. Liu 952223 (MO, SHM); Guo-zi-gou, 44°21′N, 88°59′E, Whittemore et al. 4790B (MO); Qing-he Co., 46°38′N, 90°25′E, Whittemore et al. 4605A (MO). Xizang: Shuang-hu Co., K.-Y. Lang 1361, 1364, 1367 (all in MO, PE), 1350 (IFSBH, PE).

Notes     (Last Modified On 5/4/2009)
general taxon notes:
Grimmia anodon is characterized by the absence of peristome teeth and by having subglobose, ventricose capsules that are immersed on very short and curved setae. This is the only species of Grimmia in China that has no peristome and is easily recognized when sporophytes are present. In sterile condition, it is recognized by rather small and slender plants, often in dense hoary cushions, and by the enlarged oblong-lanceolate upper leaves that end in long, denticulate hyaline hair-points. The hyaline hair-points from the Chinese specimens are not as well developed as in those specimens from Europe and North America. This species is closely related to G. poecilostoma Card. & Sebille; however, the latter differs in having peristome. In China, Grimmia anodon can sometimes be confused with poorly developed G. laevigata. The two species differ from each other in the following characters: Grimmia anodon has soft shiny plants, longer, thin-walled basal leaf cells, and unistratose upper leaf cells; G. laevigata has rigid and dull plants, shorter basal leaf cells with thick transverse walls, and obscure, bistratose upper leaf cells.
J.-X. Luo and P.-C. Wu (1980) described Schistidium tibetanum as having dioicous plants and falling peristome. However, our examination shows that it is autoicous with perigonia found just below perichaetia and it lacks peristome teeth. They described the short setae for this species. In fact, the setae are not only shorter than capsule urns, but also distinctly curved. The capsules are asymmetric and ventricose, which they failed to mention. All these features indicate that Schistidium tibetanum is conspecific with Grimmia anodon.


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1. Grimmia anodon Bruch & Schimp. in B.S.G.   无齿紫萼藓   wu-chi zi-e xian
Bryol. Eur. 3: 110. tab. 236 (fasc. 25–28). 1845. Anodon ventricosus Rabenh., Deutschl. Krypt.-Fl. 2(3): 154. 1848. Schistidium anodon (Bruch & Schimp.) Loeske, Laubm. Eur. Part I: 49. 1913. Gasterogrimmia anodon (Bruch & Schimp.) Buyss., Feuille Jeunes Naturalistes 13: 63. 1883. Type. Germany: “In muris et ad rupes calcareas; prope Heidelberg ad arcem (A. Braun) (lectotype BM, designated by T. Cao & Vitt 1986); in monte pinifero prope Gefrees (Funk) (syntype BM).
Schistidium tibetanum J.-X. Luo & P.-C. Wu, Acta Phytotax. Sin. 18(l): 121. 1980. Type. China: Xizang (Tibet): Ban-ge Co., B.-S. Li 7601 (holotype PE).
Plants rather small, up to 1.5 cm high, dark green to brownish above, brown to dark brown below, in dense cushions. Stems branched from innovations, with well developed central strand in cross section. Leaves erect, imbricate when dry, erect-spreading when moist, fragile, 1.2–1.9 mm long, the upper leaves enlarged, broadly oblong-lanceolate, slightly concave, acute, ending in a long, denticulate hyaline hair-point; the lower leaves oblong-lanceolate to lanceolate, blunt, muticous or short-awned; margins plane, entire; costa prominent, ending near the apex, with small median cells in cross section; upper cells unistratose or in places bistratose near or at the margins, 7–10 µm wide, rounded-quadrate, with thick, more or less nodulose walls; basal cells elongate-rectangular, pale and hyaline, with straight, thin walls. Autoicous. Perichaetial leaves similar to the upper stem leaves. Setae shorter than capsules, slender, curved; capsules immersed, small, 0.6–0.8 mm long, subglobose, ventricose, wide-mouthed after dehiscence; exothecial cells elongate with thin walls; annuli wanting; opercula plano-convex, bluntly mucronate; peristome none. Calyptrae mitrate, deeply lobed, smooth. Spores 8–11 µm in diameter, yellowish brown, granulose in regular rows.


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