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Published In: Die Natürlichen Pflanzenfamilien 229[I,3]: 1008. 1907. (Nat. Pflanzenfam.) Name publication detail

Project Name Data (Last Modified On 3/30/2009)
Acceptance : Accepted
Project Data     (Last Modified On 3/30/2009)
General Reference:

Illustration & Map     (Last Modified On 3/30/2009)
Related Objects:

Illustrations: P.-C. Wu (ed.) 2001 (Pl. 88, figs. 1–17).

Click here for distribution map.

Map: Haplocladium angustifolium (Hampe & Müll. Hal.) Broth. (Distribution in China)

Distribution     (Last Modified On 3/30/2009)
Habitat: mostly on rocks, rarely on the bases of trees or rotten logs; alt. 1500–1700 m.
Distribution: China, Korea, Japan, Vietnam, Burma, India, Pakistan, Russia (Siberia), Europe, and Africa.

Specimens Examined     (Last Modified On 3/30/2009)
Specimens Examined:
Chinese specimens examined: Chongqian: Mt. Jinfou, M.-Z. Wang 21146 (PE). Guangdong: Mt. Luofu, Magill et al. 8139 (MO). Guizhou: Gui-yang City, Gui-yang Teacher’s Coll. 139 (MO, PE); Ping-ba (Pin-fa) Co., Cavalerie 1737 (PC). Hebei: Mt. Dong-ling, L.-Q. Yu 3 (MO, PE). Henan: Xin-yang City, J.-X. Luo 75 (MO, PE). Hong Kong: Bowing s.n. (NY). Hubei: Mt. Shennongjia, P.-C. Wu 60, 61, 103, 112a (MO, PE); Xing-shan Co., J.-X. Luo 961270 (MO, PE). Jiangsu: Nan-jing (Nanking) City, Steward 943 (BM, NY); Wu Co., P.-C. Wu & Y.-X. Jin 1631 (MO, PE). Jiangxi: Nan-chang City, Hb. Mitten, Collector unknown (NY). Nei Mongol: Zha-lan-dun City, Ba-lin, X.-L. Bai 2504 (NY). Shaanxi: Mt. Qinling, P.-C. Chen et al. 22 (MO, PE); Mt. Taibai, Z.-P. Wei 5233 (MO); Tui Kio San, Giraldi 2204 (BM, PC). Shandong: Tai-shan (Mt.), C. Gao & E.-Z. Bai 34029a (MO). Shanghai: Zaukadoo, Bailey s.n. (BM, NY). Shanxi: Yuan-qu Co., Y.-J. Ling 85021 (PE); Xin-yuan Co., Y.-J. Ling 85069, 85070, 85071 (PE, SXU). Sichuan: Lei-bo Co., P.-C. Chen 5891 (MO, PE); Li Co., X.-J. Li 915 (MO, PE); Mt. Er-lang, C. Gao 18252 (MO); Yenyuen (Yan-yuan) Co., Handel-Mazzetti 2598 (BM). Taiwan: Nang-tou Co., C.-C. Chung & W. B. Schofield 807 (BCU, NY, PE); Tai-pei Co., M.-J. Lai 10053 (as Haplocladium microphyllum, MO, NY). Yunnan: An-ning Co., Magill 8062 (MO); Da-li Co., X.-Y. Jia 910075 (MO); Lu-nan Co., Redfearn et al. 2290 (MO); Meng-hai Co., Redfearn 34107 (MO); Meng-la Co., Magill et al. 7877 (MO); Rui-li Co., X.-Y. Jia 910063 (MO); Song-ming Co., Koponen 37815 (H, MO); Wei-shan Co., Redfearn et al. 1212 (MO, NY); Yang-bi Co., Redfearn & Y.-G. Su 144 (MO). Zhejiang: Hang-zhou City, Buck 23853 (NY); Sui-chang Co., Z.-L. Liu & R.-L. Hong 35 (HSNU, MO).

Notes     (Last Modified On 3/30/2009)
general taxon notes:
This species is characterized by having projecting leaf cells and long lanceolate leaf apices.


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1. Haplocladium angustifolium (Hampe & C. Müll.) Broth. 狭叶小羽藓 xia-ye xiao-yu xian
Nat. Pflanzenfam. I(3): 1008. 1907. Bryohaplocladium angustifolium (Hampe & C. Müll.) Watanabe & Iwats., J. Jap. Bot. 56: 259. 1981. Hypnum angustifolium Hampe & C. Müll., Bot. Zeitung (Berlin) 13: 88. 1855. Type. South Africa: Krakakamma, Ecklon s.n.
Haplocladium incurvum Broth., Symb. Sin. 4: 99. 1929. Type. China: Hunan, Changsha (Tschangscha) City, Handel-Mazzetti 12912 (isotype BM).
Haplocladium lutescens (Card.) Broth. ex Thér., Ann. Cryptog. Exot. 3: 82. 1930. Pseudoleskea lutescens Card., Bull. Soc. Bot. Genève, ser. 2, 3: 284. 1911. Type. China: Taiwan, Takao, Faurie 2399.
Haplocladium subulaceum (Mitt.) Broth., Nat. Pflanzenfam. I(3): 1008. 1907. Leskea subulacea Mitt., J. Proc. Linn. Soc., Bot., Suppl. 1: 131. 1859.
Haplocladium subulaceum (Mitt.) Broth. var. amblystegioides (Broth. & Par.) Thér., Ann. Cryptog. Exot. 3: 93. 1930. Haplocladium amblystegioides (Broth. & Par.) Broth., Nat. Pflanzenfam. I(3): 1008. 1917. Thuidium amblystegioides Broth. & Par., Rev. Bryol. 31: 57. 1904.
Haplocladium subulaceum (Mitt.) Broth. var. fuscissimum (C. Müll.) Thér., Ann. Cryptog. Exot. 3: 93. 1930. Thuidium fuscissimum (C. Müll.) Par., Ind. Bryol. 1281. 1898. Haplocladium fuscissimum C. Müll., Nuovo Giorn. Bot. Ital., n. ser. 4: 275. 1897. Type. China: Shaanxi (Schen-si), in monte Lao-y-san, March 1896, Giraldi s.n.
Haplocladium subulaceum (Mitt.) Broth. var. macropilum (C. Müll.) Thér., Ann. Cryptog. Exot. 3: 91. 1930. Pseudoleskea macropilum (C. Müll.) Salmon., J. Linn. Soc. Bot. 34: 471. 1900. Thuidium macropilum (C. Müll.) Par., Ind. Bryol. 1285. 1898. Haplocladium macropilum C. Müll., Nuovo Giorn. Bot. Ital., n. ser. 3: 116. 1896. Syntypes. China: Shaanxi (Schen-si), Hua-tzo-pin, 20 June 1894, Giraldi s.n.; Si-ku-tziu-san, July 1894, Giraldi s.n.
Haplocladium subulaceum (Mitt.) Broth. var. subulatum (Card.) Thér., Ann. Cryptog. Exot. 3: 91. 1930. Haplocladium subulatum Card., Bull. Soc. Bot. Genève, ser. 2, 3: 282. 1911.
Thuidium rubicundulum (C. Müll.) Par., Ind. Bryol. Suppl. 1: 321, 1900. Haplocladium rubicundulum C. Müll., Nuovo Giorn. Bot. Ital., n. ser. 5: 208. 1898. Syntypes. China: Shaanxi (Schen-si), Sche-kin-tsuen, April 1896, Giraldi 2211 (syntype BM); Huo-kia-zaea, March 1897, Giraldi s.n.; Schan-gen-ze, March 1897, Giraldi s.n.
Plants small to medium-sized, green to yellowish green, brown when old, forming interwoven patches. Stems creeping, up to 10 mm long, regularly pinnately branched, branches ca. 5 mm long; central strand present; paraphyllia lanceolate, mostly on stems and rarely on branches. Stem leaves concave, gradually tapering from ovate to widely ovate base to a slender lanceolate apex; leaf margins revolute or plane, serrate; costa stout, often excurrent; median leaf cells quadrate to rhomboidal, 5–25 µm wide, thick-walled, projecting at upper ends; apical leaf cells narrow, rhomboidal, smooth; branch leaves ovate to narrowly ovate at base, with a short or long lanceolate apex. Dioicous. Perichaetial leaves entire at the margin. Setae 3–4 cm long, orange-red; opercula conical; peristome similar to H. microphyllum (Hedw.) Broth.


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