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Published In: Die natürlichen Pflanzenfamilien, Zweite Auflage 11: 487. 1925. (Nat. Pflanzenfam. (ed. 2)) Name publication detail

Project Name Data (Last Modified On 4/3/2009)
Acceptance : Accepted
Project Data     (Last Modified On 4/3/2009)
General Reference:

Illustration & Map     (Last Modified On 4/3/2009)
Related Objects:
Illustrations: Noguchi 1994, fig. 528, B.

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Map: Hylocomiastrum umbratum (Hedw.) M. Fleisch. ex Broth. (Distribution in China)

Distribution     (Last Modified On 4/3/2009)
Habitat: mostly on moist forest floor.
Distribution: China, Japan, Korea, Russia, Europe, North America, and North Africa.

Specimens Examined     (Last Modified On 4/3/2009)
Specimens Examined:
Chinese specimens examined: Jilin: Chang-bai Co., C. Gao 7434 (MO); Mt. Changbai, C. Gao 1064, 1094, 1106 (all in IFSBH). Heilongjiang: Ning-an Co., X.-L. Zhu 241 (PE). Sichuan: Mt. Erlang, C. Gao et al. 17695 (IFSBH, PE).


Notes     (Last Modified On 4/3/2009)
general taxon notes:


This species is characterized by having double or forked costae. It is related to Hylocomiastrumhimalayanum, but differs by having bipinnately branched stems; the latter usually has tri-pinnately branched stems. Hylocomiastrumhimalayanum is endemic to Eastern Asia, from the Himalayas to Japan, while H. umbratum is more widely distributed.


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3.  Hylocomiastrum umbratum (Hedw.) Fleisch. in Broth.   仰叶星塔藓   yang ye xing ta xian
Nat. Pflanzenfam. (ed. 2), 11: 487. 1925. Hylocomium umbratum (Hedw.) Schimp. in B.S.G., Bryol. Eur. 5: 175 (fasc. 49–51 Mon. 6). 1852. Hypnum umbratum Ehrh. ex Hedw., Sp. Musc. Frond. 263. 1801. Type: Europe.
Plants slender, dark green to brownish green, slightly glossy. Main stems up to 10 cm long, densely pinnately branched above, loosely pinnately branched below; branches ca. 1.5 cm long, attenuate at apex, twisted when dry, arcuate when moist; pseudoparaphyllia lanceolate. Stem leaves triangularly lanceolate, with a piliform or shortly acuminate apex, ca. 1.8 mm × 1.2 mm, deeply plicate at base; margins somewhat reflexed at the middle, serrulate or nearly entire; costae double, often reaching the middle part of lamina; branch leaves broadly ovate, acute, ca. 1.0 mm × 0.7 mm, deeply plicate; margins serrate; costae long and strong; median cells of stem leaves linear, 40–65 µm × 4–5 µm, thin-walled, not clearly pitted; basal cells nearly linear or rectangular, walls yellowish brown, not clearly pitted; alar cells similar to basal cells, but slightly larger. Inner perichaetial leaves lanceolate, up to 1.7 mm long, entire, ecostate; paraphyllia few. Setae elongate, up to 4.0 cm long, thick, twisted when dry, curved when moist; capsules inclined, ovoid, ca. 2.5 mm × 1.5 mm; opercula conic, with an apiculate tip, ca. 1 mm long; exostome teeth ca. 0.75 mm long, yellowish brown below; endostome segments as long as the teeth. Spores 12–16 µm in diameter.


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