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Published In: The Bryologist 74(2): 167. 1971. (Bryologist) Name publication detail

Project Name Data (Last Modified On 4/2/2009)
Acceptance : Accepted
Project Data     (Last Modified On 4/2/2009)
General Reference:

Illustration & Map     (Last Modified On 4/2/2009)
Related Objects:
Illustrations: P.-C. Chen et al. 1978, pl. 351, figs. 1–10.

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Map: Callicladium haldanianum (Grev.) H.A. Crum (Distribution in China)

Distribution     (Last Modified On 4/2/2009)
Habitat: on rotten wood, base of trees or rocks; alt. 750–1700 m.
Distribution: China, Japan, Korea, Russia, Europe, and North America.

Specimens Examined     (Last Modified On 4/2/2009)
Specimens Examined:


Chinese specimens examined: Heilongjiang: Bo-li Co., Y.-L. Zhang 2155 (IFSBH, MO); Shang-zhi Co., S.-E. Liu 7938 (IFSBH, MO); Xiao Xing An Ling (Lesser Khingan Mt.), C. Gao 8, 622, 625 (all in IFSBH, MO), Dailing Distr., Z.-H. Zhang 4138 (IFSBH, MO), Shihezi Distr., C. Gao 157, 190 (both in IFSBH, MO), Wuying Distr., C. Gao 442 (IFSBH, MO), P.-C. Chen & C. Gao 421 (IFSBH, MO). Jilin: An-tu Co., C. Gao 1170, 22185 (both in IFSBH, MO); Chang-bai Co., C. Gao 7202, 7266, 22238 (all in IFSBH, MO), C. Gao & G.-C. Zhang 1144 (IFSBH, MO); Mt. Changbai, Vitt 34740 (ALTA, MO), C. Gao 22244 (IFSBH, MO); Lin-jiang Co., S.-E. Liu 1039 (IFSBH, MO). Liaoning: Kuan-dian Co., X.-L. Chen 004203 (IFSBH, MO), T. Cao & S.-Z. Wang 38458 (IFSBH, MO). Xinjiang: He-jing Co., S. He 951188 (MO). Yunnan: Da-li Co., Redfearn et al. 1471, 1544 (both in MO, SMS).

Notes     (Last Modified On 4/2/2009)
general taxon notes:
Callicladium haldanianum was recognized as a species of Heterophyllium in the Sematophyllaceae by some earlier Chinese moss floras (C. Gao 1977, P.-C. Chen et al. 1978). It was recently treated in the Hypnaceae by X.-L. Bai (1997). However, it was not treated in the most recent work, “Flora Bryophytorum Sinicorum,” neither in Hypnaceae nor in Sematophyllaceae (P.-C. Wu and Y. Jia 2004).


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1. Callicladium haldanianum (Grev.) Crum   拟腐木藓   ni fu mu xian
4: 1177. 1923. Hypnum haldanianum Grev., Ann. Lyceum Nat. Hist. New York. 1: 275. f. 23. 1825. Type: Canada.
Callicladium haldanianum (Grev.) Crum var. viride (Okam.) Iwats. & Nog., J. Jap. Bot. 48: 215. 1973. Stereodon haldanianus var. viridis Okam., J. Coll. Sci. Imp. Univ. Tokyo, 36(7): 34. pl. 17. 1915.
Plants medium-sized, bright-green to yellowish or brownish, in extensive, flattened mats. Stems creeping, irregularly branched; branches rather short and unequal, mostly flattened, tapered; pseudoparaphyllia foliose. Stem leaves and branch leaves similar, crowded, erect to erect-spreading, usually homomallous, somewhat complanate, concave, clearly asymmetric, more or less keeled, ovate-lanceolate, gradually narrowed to a short to rather long acumen, 1.0–1.6 mm × 0.5–0.6 mm; margins plane, entire; costa very short, double or sometimes indistinct; median and upper leaf cells linear, smooth, flexuose; lower cells shorter and broader; alar cells clearly differentiated, excavate, usually yellowish brown, quadrate to subquadrate, moderately thick-walled. Autoicous. Perichaetial leaves long-subulate and sheathing at base. Setae elongate, 1.5–3.0 cm long, smooth; capsules long-cylindrical, 1.5–3.0 mm long, asymmetric, suberect from the base and somewhat curved near the mouth; annuli slightly differentiated; opercula shortly obliquely rostrate; peristome double; exostome teeth narrowly lanceolate, cross-striolate below, bordered, trabeculate on the ventral surface; endostome segments slenderly lanceolate, keeled; basal membrane high; cilia single. Calyptrae cucullate, smooth. Spores spherical, 12–15 µm in diameter, finely papillose.


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